ghilzai
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Here is a brief review
Actually there is no original homeland but homelands as the Pashtun evolved from many different people
the basic Ones are east Iranic tribes and later the mixing of Scythian tribes(For ex the Sarakzai claim descent from Scythians) , there is a theory that some of the lost tribes of Judea mixed came to Afghanistan and beyond and there is some basic proof although there are still holes to be solved in that theory(Yosufzai is the largest pashtun tribe , Yousafzai stands for son of Yousuf/Joseph) Then there were the Turkic people , alot of the tribes pashtunized most famous being the Ghilzais!
There is also a theory that Kashmiris were basically the same people as Pashtuns who ventured into the Kashmir valley and were baffled by its beauty and never returned. Later the diffrences arised because both mixed with different people and were influenced by different groups/civilization!
I would have given you a better explaination with appropriate links but I am extremely busy these days due to exams.
I had an extremely good link with refrences given about the lost tribe theory, if I find it i'l share it.
Yusufzais are not Semitic this was a label applied to make them sound Bani Israel and thus ensuring automatic right to rule.
They are also not the biggest tribe, they are mostly farmers that have been pushed towards the Indus River.
Pukhtoons can be iranic or Turkic, the only thing is some of us will never use the word afghan as to us it's always khurasan.
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